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StableTalk
- The UK's brightest on-line equestrian magazine, written
by riders for riders
The
Ramp Retina - What is it and does it exist
Ref: AM Harman, S Moore, R Hoskins and P Keller
It
is commonly observed that when a horse looks at something
in the distance it raises its head and lifts its nose and
when observing much closer objects the horse may arch the
neck and rotate the head on one side. In the past it has
been explained that the horse needs to make up for an inability
to focus on objects when viewing at different distances
(accommodation) by using a ramp retina. This
concept was used to denote that there is a variable distance
from the lens to the retina (the deepest part of the eye)
in the eye of the horse. It was assumed that this retinal
feature accounted for the alterations in head position exhibited
by horses. The horse was presumed to be changing the angle
of the head in order to find a visual axis along the ramp
retina with a distance from the lens to retina appropriate
for focussing on the image.
Researchers from Western Australia have conducted a study
to examine the eye of the horse to confirm or deny the presence
of a ramp retina. Other aims of the study were
to discover the anatomical features that might account for
the head moving behaviour, to confirm whether horses have
limited accommodation as well as to estimate the visual
field of the horse.
Results
from this study show that the horse does not have a ramp
retina. It was found that the horse has a retina that
is the same distance from the lens in all areas except in
the peripheral regions where the retina is closer to the
lens. This is opposite to previous theories of the ramp
retina. The study also found that the horse can see
a very narrow but panoramic view clearly, and this area
of vision is perceived by a retinal region called the visual
streak. The horses peripheral vision is poor; a ramp
retina would be of little benefit to the horse, as
it would give the horse no advantage to use any part of
the retina other than the visual streak for direct observation.
Results
from the visual field tests show that the binocular overlap
(areas where the horse is viewing objects with both eyes
at once) is down the nose and not straight-ahead. In fact
the horse has a blind spot straight in front of the forehead.
Therefore the horse has to raise its nose considerably to
see directly in front. When the horse lowers its head, the
binocular field is directed towards the ground for grazing
and the monocular fields are in position to scan the lateral
horizon. When the head is raised with the nose pointing
forward, the horse uses the binocular field (both eyes)
to scan the horizon and the monocular (single eye) lateral
vision becomes limited. The monocular fields are used during
calm moments of grazing when the head is in the position
that the nose approaches the vertical and it is this head
orientation that many riders attempt to obtain from their
mounts when their horse is on the bit and which
results in relaxation and submission from the horse. When
the horse is distracted by a sound or sees something in
its peripheral vision, it will turn its head or whole body
towards the object and raise its head. The researchers believe
the head is raised so the horse can utilise its binocular
vision to see something in the distance and not to alter
the focussing power of the eye as suggested by the concept
of the ramp retina. When the horse is looking
at something close, for example a treat held in the hand,
the researchers believe that the horse arches its neck and
rotates its head in order to utilise the binocular vision
down the nose.
Another
important finding is that the researchers believe the blind
spot in front of the horse is actually the width of the
horse and when the horse is on the bit it cannot
see anything directly in front of it and must rely on the
rider. Therefore a fair degree of submission is requested
by the rider and given by the horse to achieve this position.
If a showjumping horse is to see and judge the distance
of a fence that it approaches, it must be allowed the freedom
to raise its head and direct its binocular field forward.
Click
here for photos that try to represent the difference
between equine and human vision.
Photo Information
The view seen by the left eye of a human and
a horse facing the same direction, looking over
the city of Perth (Australia). (A) is the front view
of the city and (B) is the rear view behind the horse
and the human. (C) is the view seen by the human,
a small high acuity central region surrounded by a
lower acuity area and (D) is the view seen by the
horse, running from the city almost to the rear as a
horizontal strip with much lower acuity above and below.
Photo courtesy of the Equine Veterinary Journal.
Click
here for the transcript of a report featured on ABC
television in Australia.
Ref:
AM Harman, S Moore, R Hoskins and P Keller Horse Vision
and an Explanation for the Visual Behaviour Originally explained
by the Ramp Retina (1999) Equine Veterinary
Journal Vol. 31, No. 5, Pg. 384.
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